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This chapter is an overview of the structure of words belonging to the major lexical categories (nouns and verbs) in Niger-Congo languages, with an emphasis on the morphological patterns typically found in the core Niger-Congo languages commonly considered as relatively conservative in their morphology: rich systems of verb morphology, both inflectional and derivational, and systems of gender-number marking with a relative high number of genders, and no possibility to isolate number marking from gender marking. As regards formal aspects of the structure of words, as a rule, verb forms are morphologically more complex than nominal forms. The highest degree of synthesis is found in the verbal morphology of some Bantu languages. Both prefixes and suffixes are found. Cumulative exponence is typically found in gender-number marking. Multiple exponence is very common in the verbal morphology of Bantu language but rather uncommon in the remainder of Niger-Congo. Consonant alternations are common in several groups of Niger-Congo languages, and various types of tonal alternations play an important role in the morphology of many Niger-Congo languages. The categories most commonly expressed in the inflectional morphology of nouns are gender, number, definiteness, and possession. The inflectional morphology of verbs commonly expresses agreement, TAM, and polarity, and is also widely used to express interclausal dependencies and information structure. As regards word formation, the situation is not uniform across the language groups included in Niger-Congo, but rich systems of verb-to-verb derivation are typically found in the Niger-Congo languages whose morphological patterns are commonly viewed as conservative.

The category of Personal/Participant/Inhabitant derived nouns comprises a conglomeration of derived nouns that denote among others agents, instruments, patients/themes, inhabitants, and followers of a person. Based on the thematic relations between the derived noun and its base lexeme, Personal/Participant/Inhabitant nouns can be classified into two subclasses. The first subclass comprises derived nouns that are deverbal and carry thematic readings (e.g., driver). The second subclass consists of derived nouns with athematic readings (e.g., Marxist). The examination of the category of Personal/Participant/Inhabitant nouns allows one to delve deeply into the study of multiplicity of meaning in word formation and the factors that bear on the readings of derived words. These factors range from the historical mechanisms that lead to multiplicity of meaning and the lexical-semantic properties of the bases that derived nouns are based on, to the syntactic context into which derived nouns occur, and the pragmatic-encyclopedic facets of both the base and the derived lexeme.

In linguistics, the study of quantity is concerned with the behavior of expressions that refer to amounts in terms of the internal structure of objects and events, their spatial or temporal extension (as duration and boundedness), their qualifying properties, as well as how these aspects interact with each other and other linguistic phenomena. Quantity is primarily manifest in language for the lexical categories of noun, verb, and adjective/ adverb. For instance, the distinction between mass and count nouns is essentially quantitative: it indicates how nominal denotation is quantized—as substance (e.g., water, sand) or as an atomic individual (e.g., book, boy). Similarly, the aspectual classes of verbs, such as states (know), activities (run), accomplishments (drown), achievements (notice), and semelfactives (knock) represent quantitatively different types of events. Adjectives and adverbs may lexically express quantities in relation to individuals, respectively, events (e.g., little, enough, much, often), and one might argue that numerals (two, twenty) are intrinsic quantitative expressions. Quantitative derivation refers to the use of derivational affixes to encode quantity in language. For instance, the English suffix -ful attaches to a noun N1 to derive another noun N2, such that N2 denotes the quantity that fits in the container denoted by N1. N2 also employs a special use in quantitative constructions: see hand—a handful of berries. The challenge for the linguistic description of quantity is that it often combines with other linguistic notions such as evaluation, intensification, quality, and it does not have a specific unitary realization—it is usually auxiliary on other more established notions. Quantitative affixes either have limited productivity or their primary use is for other semantic notions. For instance, the German suffix ‑schaft typically forms abstract nouns as in Vaterschaft ‘fatherhood’, but has a (quantity-related) collective meaning in Lehrerschaft ‘lecturer staff’; compare English -hood in childhood and the collective neighborhood. This diversity makes quantity difficult to capture systematically, in spite of its pervasiveness as a semantic notion.

In Germanic languages, conversion is seen as a change in category (i.e., syntactic category, word class, part of speech) without (overt) affixation. Conversion is attested in all Germanic languages. The definition of conversion as transposition or as derivation with a so-called zero-affix, which is responsible for the word-class change, depends on the language-specific part-of-speech system as well as, as often argued, the direction of conversion. Different types of conversion (e.g., from adjective to noun) are attested in Germanic languages, which differ especially semantically from each other. Although minor conversion types are attested, the main conversion types in Germanic languages are verb-to-noun conversion (deverbal nouns), adjective-to-noun conversion (deadjectival nouns), and noun-to-verb conversion (denominal verbs). Due to the characteristics of word-class change, conversion displays many parallels to derivational processes such as the directionality of category change and the preservation of lexical and grammatical properties of the underlying stem such as argument structure. Some, however, have argued that conversion does not exist as a specific rule and is only a symptom of lexical relisting. Another question is whether two such words are related by a conversion process that is still productive or are lexically listed relics of a now unproductive process. Furthermore, the direction of conversion of present-day Germanic, for example, the identification of the word class of the input before being converted, is unclear sometimes. Generally, deverbal and deadjectival nominal conversion in Germanic languages is semantically more transparent than denominal and deadjectival verbal conversion: despite the occurrence of some highly frequent, but lexicalized counterexamples, the semantic impact of conversion is only sometimes predictable, slightly more in the nominal domain than in the verbal domain. The semantics of verb formation by conversion (e.g., whether conversion leads to causative readings or not) is hardly predictable. Overall, conversion in Germanic is considered a process with multiple linkages to other morphological phenomena such as derivation, back-formation, and inflectional categories such as grammatical gender. Due to the lack of formal markers, conversion is considered non-iconic. The different kinds of conversions are merely based on language-specific mechanisms, but what all Germanic languages share at least is the ability to form nominal conversion, which is independent of their typological characteristics as isolating-analytic versus inflectional-fusional languages. This is surprising given the crosslinguistic prevalence of verbal conversion in the languages of the world.

Conversion is traditionally viewed as a word-formation technique of forming a word from a formally identical but categorically different word without adding a(n explicit) morphological exponent. Despite its apparent formal simplicity manifested first of all in the sameness of the input and the output, the proper understanding of what exactly happens during conversion, morphosyntactically and semantically alike, is by no means an easy matter even in respect of one language, let alone languages representing different typological groups or subgroups. To determine the linguistic status of conversion and its place among other types of word formation is not a simple matter either, and, paradoxically, it is especially so in the case of the most extensively studied English conversion. The reason for this is that the traditional view of conversion has often been called into question, giving rise to a diversity of interpretations of conversion not only in English but also in a cross-linguistic perspective. Conversion research has gone a long way to explore the mechanism of conversion as a kind of word formation; nevertheless, further research is necessary to understand every detail of this mechanism.

Denominal verbs are verbs formed from nouns by means of various word-formation processes such as derivation, conversion, or less common mechanisms like reduplication, change of pitch, or root and pattern. Because their well-formedness is determined by morphosyntactic, phonological, and semantic constraints, they have been analyzed from a variety of lexicalist and non-lexicalist perspectives, including Optimality Theory, Lexical Semantics, Cognitive Grammar, Onomasiology, and Neo-Construction Grammar. Independently of their structural shape, denominal verbs have in common that they denote events in which the referents of their base nouns (e.g., computer in the case of computerize) participate in a non-arbitrary way. While traditional labels like ‘ornative’, ‘privative’, ‘locative’, ‘instrumental’ and the like allow for a preliminary classification of denominal verbs, a more formal description has to account for at least three basic aspects, namely (1) competition among functionally similar word-formation patterns, (2) the polysemy of affixes, which precludes a neat one-to-one relation between derivatives displaying a particular affix and a particular semantic class, and (3) the relevance of generic knowledge and contextual information for the interpretation of (innovative) denominal verbs.

Headedness is a pervasive phenomenon throughout different components of the grammar, which fundamentally encodes an asymmetry between two or more items, such that one is in some sense more important than the other(s). In phonology for instance, the nucleus is the head of the syllable, and not the onset or the coda, whereas in syntax, the verb is the head of a verb phrase, rather than any complements or specifiers that it combines with. It makes sense, then, to question whether the notion of headedness applies to the morphology as well; specifically, do words—complex or simplex—have heads that determine the properties of the word as a whole? Intuitively it makes sense that words have heads: a noun that is derived from an adjective like redness can function only as a noun, and the presence of red in the structure does not confer on the whole form the ability to function as an adjective as well. However, this question is a complex one for a variety of reasons. While it seems clear for some phenomena such as category determination that words have heads, there is a lot of evidence to suggest that the properties of complex words are not all derived from one morpheme, but rather that the features are gathered from potentially numerous morphemes within the same word. Furthermore, properties that characterize heads compared to dependents, particularly based on syntactic behavior, do not unambigously pick out a single element, but the tests applied to morphology at times pick out affixes, and at times pick out bases as the head of the whole word.

The term productivity most commonly applies to word formation, but in principle it can be applied to any linguistic process. A process is said to be productive if it applies without restriction to give rise to novel expressions, for instance, new words. Ideally, speakers apply productive processes without conscious deliberation and in principle they can be applied by any and all competent native speakers. Creative coining of words, by contrast, is typically a one-off intentional act (nonce word creation), often of one individual, and often with the aim of drawing attention to the coined word for the purposes of advertising, humor, and so on. Creative coining often involves extra-grammatical processes which cannot be described deterministically, unlike bona fide linguistic rules. However, many productive processes are restricted in various ways, and some of the extra-grammatical devices are very frequent, so the distinction between productive, grammatical word formation and (nonce) word creation is blurred.

Evaluative morphology is a field of linguistic studies that deals with the formation of diminutives, augmentatives, pejoratives, and amelioratives. Actually, evaluative constructions cross the boundaries of morphology, and are sometimes realized by formal strategies that cannot be numbered among word formation processes. Nevertheless, morphology plays a dominant role in the formation of evaluatives. The first attempt to draw an exhaustive account of this set of complex forms is found in the 1984 work Generative Morphology, by Sergio Scalise, who made the hypothesis that evaluatives represent a separate block of rules between inflection and derivation. This hypothesis is based on the fact that evaluatives show some properties that are derivational, others that are inflectional, and some specific properties that are neither derivational nor inflectional. After Scalise’s proposal, almost all scholars have tried to answer the question concerning the place of evaluative rules within the morphological component. What data reveal is that, in a cross-linguistic perspective, evaluatives display a uniform behavior from a semantic and functional point of view, but exhibit a wide range of formal properties. In other words, functional identity does not imply formal identity; consequently, we can expect that constructions performing the same function display different formal properties in different languages. So, if evaluatives are undoubtedly derivational in most Indo-European languages (even if they cannot be considered a typical example of derivation), they are certainly quite close to inflection in some Bantu languages. This means that the question about the place of evaluatives within the morphological component probably is not as crucial as scholars have thought, and that other issues, sometimes neglected in the literature, deserve the same attention. Among them, the role of pragmatics in the description of evaluatives is no doubt central. According to Dressler and Merlini Barbaresi, in their 1994 work, Morphopragmatics: Diminutives and Intensifiers in Italian, German and Other Languages, evaluative constructions are the more typical instantiation of morphopragmatics, which is “defined as the area of general pragmatic meanings of morphological rules, that is of the regular pragmatic effects produced when moving from the input to the output of a morphological rule.” Evaluatives include “a pragmatic variable which cannot be suppressed in the description of [their] meaning.” Another central issue in studies on evaluative morphology is the wide set of semantic nuances that usually accompany diminutives, augmentatives, pejoratives, and amelioratives. For example, a diminutive form can occasionally assume a value that is attenuative, singulative, partitive, appreciative, affectionate, etc. This cluster of semantic values has often increased the idea that evaluatives are irregular in nature and that they irremediably avoid any generalization. Dan Jurafsky showed, in 1996, that these different meanings are often the outcome of regular and cross-linguistically recurrent semantic processes, both in a synchronic and in a diachronic perspective.

An agent noun is a derived noun whose general meaning is ‘person who does . . .’. It is thus characterized by the feature [+ Human], regardless of whether the person involved actually performs an action (e.g., French nageur ‘swimmer’, i.e., ‘a person who swims’), carries out a profession (e.g., Spanish cabrero ‘goatherd’, i.e., ‘a person who looks after goats’), adheres to a certain ideology or group (e.g., Italian femminista ‘feminist’, i.e., ‘a person who supports or follows the feminist movement’), and so on. Agent nouns are for the most part denominal (as with cabrero and femminista above) and deverbal (as with nageur above). Latin denominal agent nouns were mainly formed with -arius, though the Latin agentive suffix par excellence was -tor, which derived nouns from verbs. Latin denominal agents were also formed with -ista, a borrowing from Greek -ιστήϛ. The reflexes of all three suffixes are widespread and highly productive in the Romance languages, as in the case of Portuguese/Spanish/Catalan/Occitan pescador ‘fisherman’ (-dor < -torem), French boucher ‘butcher’ (-er < -arium), and Romanian flautist (-ist < -ista). At any rate, the distinction between denominal and deverbal agent nouns is not always straightforward, as demonstrated by the Romance forms connected with the Latin present particle -nte, for whereas the majority display a verbal base (e.g., Italian cantante ‘singer’ ← cantare ‘to sing’), there are some which do not (e.g., Italian bracciante ‘hired hand’ ← braccio ‘arm’), thus allowing them to be regarded as denominal derivations. A minor group of agent nouns is made up of deadjectival derivations, often conveying a pejorative meaning; such is the case with Italian elegantone ‘person of overblown elegance’ (← elegante ‘elegant’) and French richard ‘very rich person’ (← riche ‘rich’).

Despite the intrinsically greater capacity for axons to regenerate in injured peripheral nerves than after injury to the central nervous system, functional recovery after most nerve injuries is very poor. A need for novel treatments that will enhance axon regeneration and improve recovery is substantial. Several such experimental treatments have been studied, each based on part of the stereotypical cellular responses that follow a nerve injury. Genetic manipulations of Schwann cells that have transformed from a myelinating to a repair phenotype that either increase their production of axon growth-promoting molecules, decrease production of inhibitors, or both result in enhanced regeneration. Local or systemic application of these molecules or small molecule mimetics of them also will promote regeneration. The success of treatments that stimulate axonal protein synthesis at the site of the nerve injury and in the growing axons, an early and important response to axon injury, is significant, as is that of manipulations of the types of immune cells that migrate into the injury site or peripheral ganglia. Modifications of the extracellular matrix through which the regenerating axons course, including the stimulation of new blood vessel formation, promotes the navigation of nascent regenerating neurites past the injury site, resulting in greater axon regeneration. Experimental induction of expression of regeneration associated gene activity in the cell bodies of the injured neurons is especially useful when regenerating axons must regenerate over long distances to reinnervate targets. The consistently most effective experimental approach to improving axon regeneration in peripheral nerves has been to increase the activity of the injured neurons, either through electrical, optical, or chemogenetic stimulation or through exercise. These activity-dependent experimental therapies show greatest promise for translation to use in patients.

Understanding of the various types of plasticity that occur in the spinal cord, as well as understanding of spinal cord functions, has vastly improved over the past 50 years, mainly due to an increase in the number of research studies and review articles on the subject. It is now understood that the spinal cord is not merely a passive conduit of neural impulses. Instead, the spinal cord can independently execute complex functions. Numerous experimental approaches have been utilized for more targeted exploration of spinal cord functions. For example, isolating the spinal cord from supraspinal influences has been used to demonstrate that simple forms of learning can be performed by spinal neuronal networks. Moreover, reduced preparations, such as acute spinal cord slices, have been used to show that spinal neurons undergo different types of modulation, including activity-dependent synaptic modification. Most spinal cord processes, ranging from integration of incoming sensory input to execution of locomotor outputs, involve plasticity. Nociceptive processing that leads to pain and spinal learning is an example of plasticity that is well-studied in the spinal cord. At the neural level, both processes involve an interplay of cellular mediators, which include glutamate receptors, protein kinases, and growth factors. The neurotrophin brain-derived neurotrophic factor (BDNF) has also been implicated in these processes, specifically as a promoter of both pro-nociception and spinal learning mechanisms. Interestingly, the role of BDNF in mediating spinal plasticity can be altered by injury. The literature spanning approximately 5 decades is reviewed and the role of BDNF is discussed in mediating cellular plasticity underlying pain processing and learning within the spinal cord.

Phonotactics is the study of restrictions on possible sound sequences in a language. In any language, some phonotactic constraints can be stated without reference to morphology, but many of the more nuanced phonotactic generalizations do make use of morphosyntactic and lexical information. At the most basic level, many languages mark edges of words in some phonological way. Different phonotactic constraints hold of sounds that belong to the same morpheme as opposed to sounds that are separated by a morpheme boundary. Different phonotactic constraints may apply to morphemes of different types (such as roots versus affixes). There are also correlations between phonotactic shapes and following certain morphosyntactic and phonological rules, which may correlate to syntactic category, declension class, or etymological origins. Approaches to the interaction between phonotactics and morphology address two questions: (1) how to account for rules that are sensitive to morpheme boundaries and structure and (2) determining the status of phonotactic constraints associated with only some morphemes. Theories differ as to how much morphological information phonology is allowed to access. In some theories of phonology, any reference to the specific identities or subclasses of morphemes would exclude a rule from the domain of phonology proper. These rules are either part of the morphology or are not given the status of a rule at all. Other theories allow the phonological grammar to refer to detailed morphological and lexical information. Depending on the theory, phonotactic differences between morphemes may receive direct explanations or be seen as the residue of historical change and not something that constitutes grammatical knowledge in the speaker’s mind.

The topic “argument structure and morphology” refers to the interaction between the number and nature of the arguments taken by a given predicate on the one hand, and the morphological makeup of that predicate on the other. This domain turns out to be crucial to the study of a number of theoretical issues, including the nature of thematic representations, the proper treatment of irregularity (both morphophonological and morphosemantic), and the very place of morphology in the architecture of the grammar. A recurring question within all existing theoretical approaches is whether word formation should be conceived of as split across two “places” in the grammar, or as taking place in only one.

Grammaticalization is traditionally defined as the gradual process whereby a lexical item becomes a grammatical item (primary grammaticalization), which may be followed by further formal and semantic reduction (secondary grammaticalization). It is a composite change that may affect both phonological, morphological, syntactic, and semantic-pragmatic properties of a morpheme, and it is found in all the world’s languages. On the level of morphology, grammaticalization has been shown to have various effects, ranging from the loss of inflection in primary grammaticalization to the development of bound morphemes or new inflectional classes in secondary grammaticalization. Well-known examples include the development of future auxiliaries from motion verbs (e.g., English to be going to), and the development of the Romance inflection future (e.g., French chanter-ai ‘I sing’, chanter-as ‘you sing’, etc., from a verb meaning ‘to have’). Although lexical-grammatical change is overwhelmingly unidirectional, shifts in the reverse direction, called degrammaticalization, have also been shown to occur. Like grammaticalization, degrammaticalization is a composite change, which is characterized by an increase in phonological and semantic substance as well as in morphosyntactic autonomy. Accordingly, the effects on morphology are different from those in grammaticalization. In primary degrammaticalization new inflections may be acquired (e.g., the Welsh verb nôl ‘to fetch,’ from an adposition meaning ‘after’), and erstwhile bound morphemes may become free morphemes (e.g., English ish). As such effects are also found in other types of changes, degrammaticalization needs to be clearly delineated from those. For example, a shift from a minor to a major category (e.g., English ifs and buts) or the lexicalization of bound affixes (isms), likewise result in new inflections, but these are instantaneous changes, not gradual ones.

Quechuan is a family of closely related indigenous languages spoken in Argentina, Bolivia, Colombia, Ecuador, and Peru, in the central part of the Andean cordilleras, in what used to be the Empire of the Incas and adjacent areas. It is divided into two main branches, commonly denominated Quechua I and II, and comprises 15 or more spoken varieties and several extinct ones that can be considered separate languages, although an exact number cannot easily be established. Quechuan shares a long and intense contact history with the neighboring Aymaran languages, but a genealogical relationship between the two families has never been demonstrated, nor a relationship with any other language family in the area. Quechuan languages are mainly agglutinative. All grammatical categories are indicated by suffixes with very few exceptions. The order in which these suffixes occur within a word form is governed by rules and combinatory restrictions that can be rigid but not always explicable on a basis of scope and function. Portmanteau suffixes play a role in verbal inflection and in mutually interrelated domains of aspect and number in the Quechua I branch. In Quechuan verbal derivation affixes may be semantically polyvalent, depending on the combinations in which they occur, pragmatic considerations, the nature of the root to which they are attached, their position in the affix order, and so on. Verbal derivational affixes often combine with specific verbal roots to denote meanings that are not fully predictable on the basis of the meaning of the components. Other verbal affixes never occur in such combinations. Verbal morphology and nominal morphology tend to overlap in the domain of personal reference, where subject and possessor markers are largely similar. Otherwise, the two morphological domains are almost completely separate. Not only the morphological inventories but also the formal constraints underlying the structure of verbs and nouns differ. Nominal expressions feature an elaborate but relatively instable system of case markers, some of which appear to be of recent formation. Transposition from one class to another, nominalization in particular, is indicated morphologically and occupies a central place in Quechuan grammar, particularly in interaction with case. Finally, there is a class of Independent suffixes that can be attached to members of all word classes, including adverbial elements that cannot be classified as verbs or nominals. These suffixes play a role at the organizational level of larger syntactic units, such as clauses, nominal phrases, and sentences.

The scope of classical generative morphology is not clearly determined. All three components need clarification. The boundaries of what counts as generative linguistics are not unambiguously set, but it can be assumed that all generative work in linguistics is inspired by the work of Noam Chomsky. Morphology was a much more prominent component of linguistic theory in earlier approaches, but of course the subject field had to be accounted for also in generative linguistics. The label classical can be seen as restricting the scope both to the more mainstream approaches and to a period that ends before the present. Here, the early 1990s will be taken as the time when classical theorizing gave way to contemporary generative morphology. In the earliest presentations of generative linguistics, there was no lexicon. The introduction of the lexicon made many of the ideas formulated before obsolete. Chomsky’s Lexicalist Hypothesis provided the basis for a new start of research in morphology. Two contrasting elaborations appeared in the early 1970s. Halle proposed a model based on the combination of morphemes, Jackendoff one based on the representation and analysis of full words. Against this background, a number of characteristic issues were discussed in the 1970s and 1980s. One such issue was the form of rules. Here there was a shift from transformations to rewrite rules. This shift can be seen particularly well in the discussion of verbal compounds, e.g., truck driver. The question whether and how morphology should be distinguished from syntax generated a lot of discussion. Another broad question was the degree to which rules of morphology should be thought of as operating in separate components. This can be observed in the issue of the distinction of inflection and derivation and in level ordering. The latter was a proposal to divide affixes into classes with different phonological and other effects on the base they attach to. A side effect of level ordering was the appearance of bracketing paradoxes, where, for instance, generative grammarian has a phonological constituent grammarian but a semantic constituent generative grammar. Another aspect of rule application which can be constructed as a difference between morphology and syntax is productivity. In general, syntactic rules are more productive and morphological rules display blocking effects, where, for instance, unpossible is blocked by the existence of impossible. Being classical, much of the discussions in this period serves as a shared background for the emergence and discussion of current generative approaches in morphology. The transition to these theories started in the 1990s, although some of them appeared only in the early 2000s.

Speakers can transfer meanings to each other because they represent them in a perceptible form. Phonology and syntactic structure are two levels of linguistic form. Morphemes are situated in-between them. Like phonemes they have a phonological component, and like syntactic structures they carry relational information. A distinction can be made between inflectional and lexical morphology. Both are devices in the service of communicative efficiency, by highlighting grammatical and semantic relations, respectively. Morphological structure has also been studied in psycholinguistics, especially by researchers who are interested in the process of visual word recognition. They found that a word is recognized more easily when it belongs to a large morphological family, which suggests that the mental lexicon is structured along morphological lines. The semantic transparency of a word’s morphological structure plays an important role. Several findings also suggest that morphology plays an important role at a pre-lexical processing level as well. It seems that morphologically complex words are subjected to a process of blind morphological decomposition before lexical access is attempted.